Species Profile and Threats Database

For information to assist proponents in referral, environmental assessments and compliance issues, refer to the Policy Statements and Guidelines (where available), the Conservation Advice (where available) or the Listing Advice (where available).
In addition, proponents and land managers should refer to the Recovery Plan (where available) or the Conservation Advice (where available) for recovery, mitigation and conservation information.

EPBC Act Listing Status Listed as Vulnerable
Listing and Conservation Advices Commonwealth Conservation Advice on Pseudocephalozia paludicola (Threatened Species Scientific Committee, 2008ez) [Conservation Advice].
Recovery Plan Decision Recovery Plan not required, included on the Not Commenced List (1/11/2009).
Adopted/Made Recovery Plans
Federal Register of
    Legislative Instruments
Declaration under s178, s181, and s183 of the Environment Protection and Biodiversity Conservation Act 1999 - List of threatened species, List of threatened ecological communities and List of threatening processes (Commonwealth of Australia, 2000) [Legislative Instrument].
State Government
    Documents and Websites
TAS:Threatened Species Notesheet - Pseudocephalozia paludicola (Tasmanian Department of Primary Industries, Parks, Water and Environment (Tas. DPIPWE), 2003n) [Information Sheet].
State Listing Status
VIC: Listed as Threatened (Flora and Fauna Guarantee Act 1988 (Victoria): February 2014 list)
Non-statutory Listing Status
VIC: Listed as Vulnerable (Advisory List of Rare or Threatened Plants in Victoria: 2005)
Scientific name Pseudocephalozia paludicola [66441]
Family Lepidoziaceae:Bryales:Bryopsida:Bryophyta:Plantae
Species author R.M.Schuster
Infraspecies author  
Reference Nova Hedwigia: Zeitschrift fur kryptogamenkunde 10:21 (1965)
Distribution map Species Distribution Map

This is an indicative distribution map of the present distribution of the species based on best available knowledge. See map caveat for more information.

Illustrations Google Images

Scientific name: Pseudocephalozia paludicola

Common name: Alpine Leafy Liverwort

Conventionally accepted as Pseudocephalozia paludicola (McCarthy 2006).

The Alpine Leafy Liverwort is a tiny, sub-alpine liverwort with creeping, often interwoven, darkish green shoots; each with three rows of leaves and leafless, pale ventral stolons (above ground runners) (Schuster & Engel 1974; Scott 1985). The leafy shoots are whitish-green to pale brown (possibly with age), soft-textured and often fleshy. The shoots form loose tufts and grow to 1–3 cm in length and 0.75–2.5 mm in width. The lateral leaves have 3–4 triangular lobes and are sharply tipped with entire margins, while the underleaves are also variably 3–4 lobed but vary from small to the same size as the lateral leaves. The leaves are concave (like cupped hands) (Scott 1985; Tas. DPIPWE 2003n). The male reproductive parts are often situated on the leafy leading stems. The main stem is generally circular or barely flattened. The female reproductive parts are short. In the field, most Pseudocephalozia species are a light, often lucid green (Scott 1985; Tas. DPIPWE 2003n).

In Tasmania, the Alpine Leafy Liverwort is known from approximately nine locations in the west, central highlands, and the eastern mountains. In Victoria, the species is known from the area around the Mount (Mt) Baw Baw plateau; namely Mt Baw Baw and Mt Kernot (MBBARMB 2007; Scott 1997; Tas. DPIPWE 2003n).

Important sites for the species in Tasmania include Olivia Plains (Cradle Valley), Mt Rufus, Shadow Lake, Hatfield (near Waratah), Leary's Corner (Middlesex Plains), Dublin Bog, south-east of Ben Nevis and the confluence of the Whyte and Pieman Rivers (Tas. DPIPWE 2003n).

In Tasmania, the species is reserved in the Central Plateau Conservation Area, Cradle Mountain-Lake St. Clair National Park, Franklin-Gordon Wild Rivers National Park, Meredith Range Regional Reserve and the Walls of Jerusalem National Park (Tas. DPIPWE 2003n). In Victoria, the Alpine Leafy Liverwort is reserved in Baw Baw National Park (Vic. DCE 1992).

The Alpine Leafy Liverwort is found in moist to wet subalpine sites, where it mostly occurs on permanently damp mineral soil or over peat. Furthermore, the species is known to occur between 1150–1500 m above sea level, in areas that are subject to snow cover for several months each year (Schuster & Engel 1974).

The species is found in moorland or sphagnum-dominated communities in Tasmania, where associated liverworts include Microlepidozia spp., Cephaloziella pulcherrima and Riccardia cochleata. The species is also occasionally associated with Eucalypt forest containing King Billy Pine (Athrotaxis selaginoides) (Schuster 2000; Schuster & Engel 1974; Tas. DPIPWE 2003n).

At the Victorian sites, the species' habitat is described as 'grassland and pools' at Mt Baw Baw. Near Mt Kernot, plants were collected on earth against a step on the Alpine Walking Track within subalpine Snow Gum (Eucalyptus pauciflora) woodland (Thies 2001 pers. comm.).

Alpine Leafy Liverwort plants reproduce by spores, and perhaps vegetatively via the stolons (above ground runners) (Sago 1998).

The main identified threats to the species include the small number and size of populations, and the apparently transient nature of the populations. Transient populations cause difficulty when relocating the species during surveys. Slope grooming and other infrastructure involved in ski field development represent potential threats to the species, where its distribution coincides with alpine resorts (Sago 2001 pers. comm.). At the Mt Kernot site within the Baw Baw National Park, the Alpine Leafy Liverwort is vulnerable to trampling by walkers or dislodgement (following heavy rain or track maintenance works) where it occurs directly on the Alpine Walking Track (Thies 2001 pers. comm.). However, it is also likely to occur away from the track in more protected sites within the Baw Baw National Park (Thies 2001 pers. comm.).

Management documents for the Alpine Leafy Liverwort can be found at the start of this profile. Other management documents relavent to the species include:

  • The Baw Baw National Park Management Plan (Vic. DCE 1992).
  • The Mount Baw Baw Alpine Resort Management Board Environmental Management Plan 2007-2010 (MBBARMB 2007).

The following table lists known and perceived threats to this species. Threats are based on the International Union for Conservation of Nature and Natural Resources (IUCN) threat classification version 1.1.

Threat Class Threatening Species References
Human Intrusions and Disturbance:Recreational Activities:Disturbance, especially from human recreational activities and development Commonwealth Conservation Advice on Pseudocephalozia paludicola (Threatened Species Scientific Committee, 2008ez) [Conservation Advice].
Species Stresses:Indirect Species Effects:Low numbers of individuals Commonwealth Conservation Advice on Pseudocephalozia paludicola (Threatened Species Scientific Committee, 2008ez) [Conservation Advice].

McCarthy, P.M. (2006). Checklist of Australian Liverworts and Hornworts. [Online]. Canberra: Australian Biological Resources Study. Available from: [Accessed: 21-May-2012].

Mount Baw Baw Alpine Resort Management Board (MBBARMB) (2007). Mount Baw Baw Alpine Resort Management Board Environmental Management Plan 2007-2010. [Online]. Available from: [Accessed: 21-May-2012].

Sago, J. (1998). The biogeography of Pseudocephalozia paludicola R. M. Schuster; an endemic Australian Liverwort. Victorian Naturalist. 115(3):84-86. Field Naturalists Club of Victoria.

Sago, J. (2001). Personal Communication.

Schuster, R.M. (2000). Beiheft 118: Austral Hepaticae Part 1. Nova Hedwigia. Berlin-Stuttgart, Germany.

Schuster, R.M. & Engel, J. (1974). A monograph of the genus Pseudocephalozia (Hepaticae). Journal of the Hattori Botanical Laboratory. 38:665-701.

Scott, G.A.M. (1985). Southern Australian Liverworts. Australian Flora & Fauna Series. 2:1-216. Canberra: AGPS.

Scott, G.A.M. (1997). Bryophytes. In: Scott G.A.M., T.J. Entwistle, T.W. May & G.N. Steven, eds. Conservation Overview of Australian Non-Marine Lichens, Bryophytes, Algae and Fungi. 99-107:23-33. Canberra: Environment Australia.

Tasmanian Department of Primary Industries, Parks, Water and Environment (Tas. DPIPWE) (2003n). Threatened Species Notesheet - Pseudocephalozia paludicola. [Online]. Available from:$FILE/Pseudocephalozia%20paludicola.pdf.

Thies, A. (2001). Personal communication.

Threatened Species Scientific Committee (2008ez). Commonwealth Conservation Advice on Pseudocephalozia paludicola. [Online]. Department of the Environment, Water, Heritage and the Arts. Available from:

Victorian Department of Conservation and Environment (Vic. DCE) (1992). The Baw Baw National Park Management Plan. [Online]. Available from: [Accessed: 19-May-2012].

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This database is designed to provide statutory, biological and ecological information on species and ecological communities, migratory species, marine species, and species and species products subject to international trade and commercial use protected under the Environment Protection and Biodiversity Conservation Act 1999 (the EPBC Act). It has been compiled from a range of sources including listing advice, recovery plans, published literature and individual experts. While reasonable efforts have been made to ensure the accuracy of the information, no guarantee is given, nor responsibility taken, by the Commonwealth for its accuracy, currency or completeness. The Commonwealth does not accept any responsibility for any loss or damage that may be occasioned directly or indirectly through the use of, or reliance on, the information contained in this database. The information contained in this database does not necessarily represent the views of the Commonwealth. This database is not intended to be a complete source of information on the matters it deals with. Individuals and organisations should consider all the available information, including that available from other sources, in deciding whether there is a need to make a referral or apply for a permit or exemption under the EPBC Act.

Citation: Department of the Environment (2014). Pseudocephalozia paludicola in Species Profile and Threats Database, Department of the Environment, Canberra. Available from: Accessed Sat, 23 Aug 2014 20:48:22 +1000.